Angel Wing Begonia losing leaves

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West Yorkshire
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United Kingdom
i have a Begonia which is the mother plant to several plants taken from cuttings. it is an awkward shape, but i figure it's been chopped about quite enough now, and it needs to rest. one of the new leaves has a hole in it, and also it's lost two leaves, one quite mature, one medium size. but it is also growing new leaves

what might be the cause of this, please? it is behind a door, so there's a bit of a draft, but i always open and shut the door as slowly as i can. could it be lack of water? in the Spring, Summer and Autumn, i've found the best way is to put them in a bucket of water for a few hours every week or so. now i've reduced the water, but maybe by too much? the moisture meter is reading very low. i suppose i was afraid of overwatering rather than underwatering.
 

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